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Faculty and Staff
XING Yaowu
Deputy Director
Academic title:
Postal Code:
Subject categories:
Biogeography and Ecology
Mailing Address:
Xishuangbanna Tropical Botanical Garden, CAS Menglun, Mengla, Yunnan 666303, China



  •   2006.9-2010.1: Ph.D. Department of Biogeography and Ecology, Kunming Institute of Botany, CAS (KIB-CAS) -The late Miocene Xianfeng flora in Yunnan, southwestern China and its paleoclimate reconstructions.  

Supervisors: Dr. Zhe-Kun Zhou (KIB-CAS).  

                     Dr. Yu-Sheng (Christopher) Liu (East Tennessee State University, U.S.A.)     

  •  2003.9-2006.7: M.Sc. Department of Biogeography and Ecology, Kunming Institute of Botany, Chinese Academy of Sciences (KIB-CAS)  -Floristic biogeography and taxonomic notes on Chinese species of Calamus (Palmae).  

        Supervisor: Prof. Kang-Lin Wang (KIB-CAS) 

  • 1999.9- 2003.7: B.Sc. Department of Agronomy, Qingdao Agricultural University, China.  



  •  2016.6-: Principal Investigator of Biogeography and Ecology Group at Xishuangbanna Tropical Botanical Garden, CAS.  
  •  2014.11-2016.5: Postdoctoral at the Field Museum of Natural History, Chicago. Work with Dr. Richard Ree. 

-Research focus: An integrative assessment of angiosperm diversification and biogeographic assembly processes in the Eastern Himalayan biodiversity hotspot. 

  • 2010.12-2014.09: Postdoctoral in University of Zurich. Work with Prof. Peter Linder 

              -Research focus: Inferring diversification patterns of angiosperms based on phylogenies and fossil record.  

  •   2010.06-2012.08: Postdoctoral in Xishuangbanna Tropical Botanical Garden. 
  •    2010.01-2010.06: Research Assistant. Department of Biogeography and Ecology, KIB-CAS. 

              -Research focus: Systematic palaeobotany and palaeoclimate reconstruction on late Cenozoic floras of S.W. China 

              -Research focus: Systematic palaeobotany and palaeoclimate reconstructions of Cenozic floras in Yunnan Province, S.W. China.  

  •  2008.12-2009.1: Short-term visiting scholar to the Institute of Palaeontology, Lanzhou University (N.W. China)  

-Advanced training on reconstructing Palaeo-CO2 concentration based on fossil cuticle analysis.  



  • Biogeographic and diversification patterns of key groups and regions. 
  •  Inferring the diversification patterns of angiosperms by using molecular and fossil data and linking the radiations to environmental changes. 
  •  Multiple-organ paleobotanical investigations providing characters of flowers, fruits, pollen, leaves and wood.  
  •   Palaeoclimate and palaeovegetation reconstructions using plant fossils.  


  Biogeography and Ecology




1.Zhang QY, Ree RH, Salamin N, Xing YW, Silvestro D: Fossil-Informed Models Reveal a Boreotropical Origin and Divergent Evolutionary Trajectories in the Walnut Family (Juglandaceae). Systematic Biology 2022, 71(1):242-258. 

2.Wu MX, Huang J, Spicer RA, Li SF, Zhao JG, Deng WYD, Ding WN, Tang H, Xing YW, Tian YM et al: The early Oligocene establishment of modern topography and plant diversity on the southeastern margin of the Tibetan Plateau. Global and Planetary Change 2022, 214. 

3.Mu QY, Yu CC, Wang Y, Han TS, Wang H, Ding WN, Zhang QY, Low SL, Zheng QJ, Peng CA et al: Comparative phylogeography of Acanthocalyx (Caprifoliaceae) reveals distinct genetic structures in the Himalaya-Hengduan Mountains. Alp Bot 2022, 132(1):153-168. 

4.Ma XD, Xing YW, Tan YH, Li JW, Zhou SS, Ding HB, Shen JY: Ceropegia jilongensis, a new species of Apocynaceae from Xizang, China. Taiwania 2022, 67(1):93-96. 

5.Zheng QJ, Yu CC, Xing YW, Han TS: A new Rorippa species (Brassicaceae), R. hengduanshanensis, from the Hengduan Mountains in China. Phytotaxa 2021, 480(3):210-222. 

6.Silvestro D, Bacon CD, Ding WN, Zhang QY, Donoghue PCJ, Antonelli A, Xing YW: Fossil data support a pre-Cretaceous origin of flowering plants. Nature Ecology & Evolution 2021, 5(4):449-+. 

7.Peng C, Yu CC, Xing YW: The complete chloroplast genome of Androsace erecta (Primulaceae) and its phylogenetic implication. Mitochondrial DNA Part B-Resources 2021, 6(7):1987-1989. 

8.Low SL, Yu CC, Ooi IH, Eiadthong W, Galloway A, Zhou ZK, Xing YW: Extensive Miocene speciation in and out of Indochina: The biogeographic history of Typhonium sensu stricto (Araceae) and its implication for the assembly of Indochina flora. Journal of Systematics and Evolution 2021, 59(3):419-428. 

9.Averyanova A, Tarasevich V, Popova S, Utescher T, Li SF, Mosbrugger V, Xing YW: Rupelian Kazakhstan floras in the context of early Oligocene climate and vegetation in Central Asia. Terra Nova 2021, 33(4):383-399. 

10.Su T, Spicer RA, Wu FX, Farnsworth A, Huang J, Del Rio C, Deng T, Ding L, Deng WYD, Huang YJ et al: A Middle Eocene lowland humid subtropical "Shangri-La" ecosystem in central Tibet. Proceedings of the National Academy of Sciences of the United States of America 2020, 117(52):32989-32995. 

11.Mu QY, Yu CC, Xing YW: Notes on the type specimen of Acanthocalyx delavayi (Caprifoliaceae) at Herbarium of the National Museum of Natural History in Paris (P). Phytotaxa 2020, 451(1):90-92. 

12.Mu QY, Yu CC, Xing YW: Notes on the type specimen of Acanthocalyx delavayi (Caprifoliaceae) at Herbarium of the National Museum of Natural History in Paris (P) (vol 451, pg 90, 2020). Phytotaxa 2020, 464(1):116-116. 

13.Low SL, Su T, Spicer TEV, Wu FX, Deng T, Xing YW, Zhou ZK: Oligocene Limnobiophyllum (Araceae) from the central Tibetan Plateau and its evolutionary and palaeoenvironmental implications. Journal of Systematic Palaeontology 2020, 18(5):415-431. 

14.Han TS, Zheng QJ, Onstein RE, Rojas-Andres BM, Hauenschild F, Muellner-Riehl AN, Xing YW: Polyploidy promotes species diversification of Allium through ecological shifts. New Phytologist 2020, 225(1):571-583. 

15.Ding WN, Ree RH, Spicer RA, Xing YW: Ancient orogenic and monsoon-driven assembly of the world's richest temperate alpine flora. Science 2020, 369(6503):578-+. 

16.Wang L, Kunzmann L, Su T, Xing YW, Zhang ST, Wang YQ, Zhou ZK: The disappearance of Metasequoia (Cupressaceae) after the middle Miocene in Yunnan, Southwest China: Evidences for evolutionary stasis and intensification of the Asian monsoon. Review of Palaeobotany and Palynology 2019, 264:64-74. 

17.Svetlana P, Torsten U, Anna A, Valentina T, Polina T, Xing YW: Early Miocene flora of central Kazakhstan (Turgai Plateau) and its paleoenvironmental implications. Plant Diversity 2019, 41(3):183-197. 

18.Su T, Farnsworth A, Spicer RA, Huang J, Wus FX, Liu J, Li SF, Xing YW, Huang YJ, Deng WYD et al: No high Tibetan Plateau until the Neogene. Science Advances 2019, 5(3). 

19.Hu JJ, Xing YW, Su T, Huang YJ, Zhou ZK: Stomatal frequency of Quercus glauca from three material sources shows the same inverse response to atmospheric pCO(2). Annals of Botany 2019, 123(7):1147-1158. 

20.Zhang QY, Huang J, Jia LB, Su T, Zhou ZK, Xing YW: Miocene Ulmus fossil fruits from Southwest China and their evolutionary and biogeographic implications. Review of Palaeobotany and Palynology 2018, 259:198-206. 

21.Li SF, Xing YW, Valdes PJ, Huang YJ, Su T, Farnsworth A, Lunt DJ, Tang H, Kennedy AT, Zhou ZK: Oligocene climate signals and forcings in Eurasia revealed by plant macrofossil and modelling results. Gondwana Research 2018, 61:115-127. 

22.Ding WN, Huang J, Su T, Xing YW, Zhou ZK: An early Oligocene occurrence of the palaeoendemic genus Dipteronia (Sapindaceae) from Southwest China. Review of Palaeobotany and Palynology 2018, 249:16-23. 

23.Xing YW, Ree RH: Uplift-driven diversification in the Hengduan Mountains, a temperate biodiversity hotspot. Proceedings of the National Academy of Sciences of the United States of America 2017, 114(17):E3444-E3451. 

24.Xing YW, Gandolfo MA, Onstein RE, Cantrill DJ, Jacobs BF, Jordan GJ, Lee DE, Popova S, Srivastava R, Su T et al: TESTING THE BIASES IN THE RICH CENOZOIC ANGIOSPERM MACROFOSSIL RECORD. International Journal of Plant Sciences 2016, 177(4):371-388. 

25.Xing YW, Gandolfo MA, Linder HP: The Cenozoic biogeographical evolution of woody angiosperms inferred from fossil distributions. Global Ecology and Biogeography 2015, 24(11):1290-1301. 

26.Schwery O, Onstein RE, Bouchenak-Khelladi Y, Xing YW, Carter RJ, Linder HP: As old as the mountains: the radiations of the Ericaceae. New Phytologist 2015, 207(2):355-367. 

27.Onstein RE, Carter RJ, Xing YW, Richardson JE, Linder HP: Do Mediterranean-type ecosystems have a common history?-Insights from the Buckthorn family (Rhamnaceae). Evolution 2015, 69(3):756-771. 

28.Jia LB, Manchester SR, Su T, Xing YW, Chen WY, Huang YJ, Zhou ZK: First occurrence of Cedrelospermum (Ulmaceae) in Asia and its biogeographic implications. Journal of Plant Research 2015, 128(5):747-761. 

29.Huang YJ, Jacques FMB, Liu YSC, Su T, Ferguson DK, Xing YW, Zhou ZK: Rubus (Rosaceae) diversity in the late Pliocene of Yunnan, southwestern China. Geobios 2015, 48(6):439-448. 

30.Hu JJ, Xing YW, Turkington R, Jacques FMB, Su T, Huang YJ, Zhou ZK: A new positive relationship between pCO(2) and stomatal frequency in Quercus guyavifolia (Fagaceae): a potential proxy for palaeo-CO2 levels. Annals of Botany 2015, 115(5):777-788. 

31.Bouchenak-Khelladi Y, Onstein RE, Xing YW, Schwery O, Linder HP: On the complexity of triggering evolutionary radiations. New Phytologist 2015, 207(2):313-326. 


33.Onstein RE, Carter RJ, Xing YW, Linder HP: Diversification rate shifts in the Cape Floristic Region: The right traits in the right place at the right time. Perspectives in Plant Ecology Evolution and Systematics 2014, 16(6):331-340. 

34.Jacques FMB, Su T, Spicer RA, Xing YW, Huang YJ, Zhou ZK: Late Miocene southwestern Chinese floristic diversity shaped by the southeastern uplift of the Tibetan Plateau. Palaeogeography Palaeoclimatology Palaeoecology 2014, 411:208-215. 

35.Hu Q, Xing YW, Hu JJ, Huang YJ, Ma HJ, Zhou ZK: Evolution of stomatal and trichome density of the Quercus delavayi complex since the late Miocene. Chinese Science Bulletin 2014, 59(3):310-319. 

36.Xing YW, Liu YSC, Su T, Jacques FMB, Zhou ZK: A new Tsuga species from the upper Miocene of Yunnan, southwestern China and its palaeogeographic significance. Palaeoworld 2013, 22(3-4):159-167. 

37.Xing YW, Hu JJ, Jacques FMB, Wang L, Su T, Huang YJ, Liu YS, Zhou ZK: A new Quercus species from the upper Miocene of southwestern China and its ecological significance. Review of Palaeobotany and Palynology 2013, 193:99-109. 

38.Wang L, Jacques FMB, Su T, Xing YW, Zhang ST, Zhou ZK: The earliest fossil bamboos of China (middle Miocene, Yunnan) and their biogeographical importance. Review of Palaeobotany and Palynology 2013, 197:253-265. 

39.Su T, Spicer RA, Liu YS, Huang YJ, Xing YW, Jacques FMB, Chen WY, Zhou ZK: Regional constraints on leaf physiognomy and precipitation regression models: a case study from China. Bulletin of Geosciences 2013, 88(3):595-608. 

40.Su T, Liu YS, Jacques FMB, Huang YJ, Xing YW, Zhou ZK: The intensification of the East Asian winter monsoon contributed to the disappearance of Cedrus (Pinaceae) in southwestern China. Quaternary Research 2013, 80(2):316-325. 

41.Su T, Jacques FMB, Spicer RA, Liu YS, Huang YJ, Xing YW, Zhou ZK: Post-Pliocene establishment of the present monsoonal climate in SW China: evidence from the late Pliocene Longmen megaflora. Climate of the Past 2013, 9(4):1911-1920. 

42.Huang YJ, Liu YS, Jacques FMB, Su T, Xing YW, Zhou ZK: First discovery of Cucubalus (Caryophyllaceae) fossil, and its biogeographical and ecological implications. Review of Palaeobotany and Palynology 2013, 190:41-47. 

43.Xing YW, Utescher T, Jacques FMB, Su T, Liu Y, Huang YJ, Zhou ZK: Paleoclimatic estimation reveals a weak winter monsoon in southwestern China during the late Miocene: Evidence from plant macrofossils. Palaeogeography Palaeoclimatology Palaeoecology 2012, 358:19-26. 

44.Huang YJ, Jacques FMB, Liu YS, Su T, Xing YW, Xiao XH, Zhou ZK: New fossil endocarps of Sambucus (Adoxaceae) from the upper Pliocene in SW China. Review of Palaeobotany and Palynology 2012, 171:152-163. 

45.Chen ST, Xing YW, Su T, Zhou ZK, Dilcher EDL, Soltis DE: Phylogeographic analysis reveals significant spatial genetic structure of Incarvillea sinensis as a product of mountain building. Bmc Plant Biology 2012, 12. 

46.Su T, Jacques FMB, Liu YS, Xiang JY, Xing YW, Huang YJ, Zhou ZK: A new Drynaria (Polypodiaceae) from the Upper Pliocene of Southwest China. Review of Palaeobotany and Palynology 2011, 164(1-2):132-142. 

47.Jacques FMB, Su T, Spicer RA, Xing YW, Huang YJ, Wang WM, Zhou ZK: Leaf physiognomy and climate: Are monsoon systems different? Global and Planetary Change 2011, 76(1-2):56-62. 

48.Jacques FMB, Guo SX, Su T, Xing YW, Huang YJ, Liu YS, Ferguson DK, Zhou ZK: Quantitative reconstruction of the Late Miocene monsoon climates of southwest China: A case study of the Lincang flora from Yunnan Province. Palaeogeography Palaeoclimatology Palaeoecology 2011, 304(3-4):318-327. 

49.Xing YW, Liu YS, Su T, Jacques FMB, Zhou ZK: Pinus prekesiya sp nov from the upper Miocene of Yunnan, southwestern China and its biogeographical implications. Review of Palaeobotany and Palynology 2010, 160(1-2):1-9. 

50.Su T, Xing YW, Liu YS, Jacques FMB, Chen WY, Huang YJ, Zhou ZK: LEAF MARGIN ANALYSIS: A NEW EQUATION FROM HUMID TO MESIC FORESTS IN CHINA. Palaios 2010, 25(3-4):234-238. 

51.Xia K, Su T, Liu YS, Xing YW, Jacques FMB, Zhou ZK: Quantitative climate reconstructions of the late Miocene Xiaolongtan megaflora from Yunnan, southwest China. Palaeogeography Palaeoclimatology Palaeoecology 2009, 276(1-4):80-86. 

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