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Faculty and Staff
PI: Community Ecology and Conservation
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Xishuangbanna Tropical Botanical Garden, the Chinese Academy of Sciences, Menglun Town, Mengla County, Yunnan, PR. China, 666303




Ph.D. (Applied Mathematics) - University of KwaZulu-Natal (1999-2006), South Africa 

M.Sc. (Environment & Development) - University of Natal (1997-1998), South Africa 

B.Sc. (Agriculture) - University of Natal (1993-1996), South Africa 

Awards: Grassland Society Medal for Best Fourth Year Project and Seminar 

Matric - Eshowe High School (1988-1992), South Africa 

Awards:   All Round Achievement; Best Actor; Public Speaking 

Koch Scholarship (3 years University Tuition) 





1. Professor (January 2016-present), Associate Professor (August 2013- December 2015) and Primary investigator (December 2013-), Community Ecology and Conservation Group, Xishuangbanna Tropical Botanical Garden (XTBG), China 


2. Postdoctoral researcher (July 2007 – July 2013) Resource Ecology Group, Wageningen University, The Netherlands   


3. Postdoctoral researcher and lecturer (August 2005 – May 2007) University of KwaZulu-Natal 



2017: Yunnan Foreign Experts Bureau Colourful Cloud Award. 

2014: Chinese Academy of Sciences: Fellowship for Young International Scientists  

1996: University of Natal: Grassland Society Medal for Best Fourth Year Project and Seminar 




Course in Advanced linear modelling, XTBG, China, 2018 - 2022 

Course in Advanced linear modelling, ATBC-AP Meeting, XTBG, China 2017. 

Basic statistics in R (AFEC-X), XTBG, China, 2013-2022. 

Course in Advanced linear modelling, ATBC, Montpellier, France, 2016. 


[During the latter part of my PhD and first postdoc I tutored and lectured undergraduate mathematics (linear algebra, calculus, and systems modelling)] 





Associate Editor: Journal of Ecology (October 2020-) 


Reviewing: Numerous journals including: Journal of Ecology, New Phytologist, Journal of Biogeography, Ecological Monographs, Biological Conservation, Journal of Applied Ecology, Transactions of the Royal Society of London B (2); Functional Ecology; Global Ecology and Conservation; Proceedings of the Royal Society London B; Perspectives in Plant Ecology, Evolution and Systematics; Global Ecology & Biogeography 




Guohualing Huang (2014) - Forest fragmentation and hunting effects on seed dispersal patterns in Xishuangbanna, China 

Carla Monoy (2015) – Temporal Changes in Tree Species and Trait Composition in a Cyclone- prone Pacific Dipterocarp Forest  

Mengqi Zhang (2018) - Investigation of herb-layer community in Yunnan savanna physiognomies and selection of species for savanna restoration  

Thiri Toe Khaing (2019) - Should Myanmar dry dipterocarp forests be called savannahs? 



Kingsly Beng (2018) - How do arthropods respond to land-use change and fragmentation in Xishuangbanna? 

Bonifacio Pasion (2018) - Plant Diversity Patterns in a Fragmented Tropical Forest Landscape in Xishuangbanna, Southwest China 

Mohamed Armani (2019) - Growth-defence strategies of spiny and non-spiny plants across the angiosperm phylogeny  

Yingying Chu (2022) - Phylogeography of two C4 grasses on the Yunnan-Guizhou Plateau  

Sedricke Lapuz (2022) - Understanding the distribution and dynamics of savannas within Tropical East Asia  

Artemis Anést (2022) - Resources and disturbances as drivers of the evolution of plant architecture 

Ninya Quibod (2023) - Morphological Diversification of the Skulls of Mammalian Herbivores under Dietary and Climatic Constraints 

Jan Sher (2023) - Kin versus non-kin performance in Chenopodium quinoa (willd) genotypes in response to different environmental factors  

Théodore Lefebvre (2023) - Ecology and evolution of spiny trunk species in regard of mammal bark feeding  



  · The role of plant architecture in plant function and resource balance. 

  · Life-history and functional diversity of savanna trees. 

  · Morphology and physiology of plants of different functional types. 

  · Distribution and evolution of anti-herbivory strategies of plants. 

  · Consequences of fragmentation for species biodiversity. 



(*corresponding author) 


1. Zhao W, Gao J, Hall D, Andersson BA, Bruxaux J, Tomlinson K, Drouzas A, Suyama Y, Wang X-R. 2024. Adaptive radiation of the Eurasian Pinus species under pervasive gene flow. New Phytologist 242: 2353-2368. http://doi.org/10.1111/nph.19694 

2. Dossa GGO, Adhikari S, …, Tomlinson KW, et al. 2024. MEETING REPORT. Lives and afterlives of lianas: 1 st International workshop on liana forest ecology, Xishuangbanna, China, 12–16 7 October 2023. New Phytologist 242: 1886-1890. 

3. Anest A, Bouchenak-Khelladi Y, Charles-Dominique T, Forest F, Caraglio Y, Hempson GP, Maurin O, Tomlinson KW*. 2024. Block then sting: the two-step evolution of defensive architecture in herbivore-driven ecosystems. Nature Plants. https://www.nature.com/articles/s41477-024-01649-4 

4. Sher J, Khan N, Tomlinson KW. 2024. Kin growth performance is more superior to non-kin under low and high- nutrient soil conditions. Plant Ecology. https://doi.org/10.1007/s11258-023-01386-2  

5. Cooper D, et al. Consistent Patterns of Common Species Across Tropical Tree Communities. Nature. https://www.nature.com/articles/s41586-023-06820-z  

6. Maurin O, Anest A, Forest F, Turner I, Barret RL, Cowan R, Wang L, Tomlinson K, Charles-Dominique T. 2023. Drift in the tropics: phylogenetics and biogeographical patterns in Combretaceae. Global Ecology & Biogeography 32: 1790-1802. https://onlinelibrary.wiley.com/doi/10.1111/geb.13737  

7. Quibod MNRM, Gelin U, van Langevelde F, Tomlinson KW*. 2023. Diet-specific responses of skull traits to moisture gradients in bovids and cervids. Linnean Society Journal for Zoological Research 199: 861-870. https://doi.org/10.1093/zoolinnean/zlad068  

8. Satdichanh M, Dossa GGO, Yan K, Tomlinson KW, Barton KE, Crow SE, Winowiecki L, V?gen T-G, Xu J, Harrison RD. Contrasting effects of plant biodiversity, aboveground biomass and edaphic factors on soil organic carbon stocks during tropical forest succession. Journal of Ecology 111: 1722-1734. https://doi.org/10.1111/1365-2745.14141 

9. dos Santos Rabelo B, van Langevelde F, Tomlinson K, Diniz P, da Silva DA, Barbosa ERM, Borghetti F. 2023. Effects of native and exotic grasses on the survival and growth of tree seedlings in a Neotropical savanna. Biological Invasions. https://doi.org/10.1007/s10530-023-03068-6  

10. Sher J, Bibi F, Jan G, Tomlinson KW, Ayaz A, Zaman W. 2023. Kin and non-kin connected plants benefit more than disconnected kin and non-kin plants under nutrient competitive environments. Plants 12: 487. https://doi.org/10.3390/plants12030487  

11. Gao J, Tomlinson KW, Wang B, Lapuz RS, Liu J-X, Pasion BO, Hai BT, Chanthayod S, Chen J, Peng Y-Q, Wang X-R. 2023. Phylogeography and introgression between Pinus kesiya and P. yunnanensis in Southeast Asia. Journal of Systematics and Evolution. https://doi.org/10.1111/jse.12949 

12. Lu Y, Zheng S, Tomlinson KW, Liu J-J. 2023. Contrasting responses of plant herbivory and disease to local and landscape drivers. Plant & Soil. https://link.springer.com/article/10.1007/s11104-023-05871-5 

13. Wang X, Fan H, Phoncharoen W, Gelin U, Tomlinson KW. 2023. Leaf chemistry of architecturally defended plants responds more strongly to soil phosphorus variation than non-architecturally defended ones. Physiologia Plantarum 175: e13856. https://doi.org/10.1111/ppl.13856 (All individual level data is available at https://doi.org/10.57760/sciencedb.07154) 

14. Yamawo A, Tomlinson KW. 2023. Defence plasticity in the spiny plant Aralia elata (Miq.) Seem. in response to light and soil fertility. Annals of Botany 131: 1073-1080. https://doi.org/10.1093/aob/mcac152 

15. Njoroge DM, Dossa GGO, Ye L, Lin X, Schaefer D, Tomlinson KW, Zuo J, Cornelissen JHC. 2023. Fauna access outweighs litter mixture effect during leaf litter decomposition. Science of the Total Environment. https://doi.org/10.1016/j.scitotenv.2022.160190 

16. Agung AP, Chornelia A, Grismer LL, Grismer JL, Quah ESH, Lu J, Tomlinson KW, Hughes AC. 2022. Description of two new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from karst landscapes in Yunnan, China highlights complex conservation needs. Zoological Research 43: 767–786. doi: 10.24272/j.issn.2095-8137.2022.105 

17. Roeder M, Dossa GGO, Cornelissen JHC, Yang X, Nakamura A, Tomlinson KW. 2022. Liana litter decomposes faster than tree litter in a multispecies and multisite experiment. Journal of Ecology 110: 2433-2447. https://doi.org/10.1111/1365-2745.13960 

18. Zhang X-W, Gelin U, …, Tomlinson KW, … et al. 2022. Rapid Eocene diversification of spiny plants in subtropical woodlands of central Tibet. Nature Communications 13: 3787. https://doi.org/10.1038/s41467-022-31512-z 

19. Bibi F, Tomlinson KW, Liu C, Liu C-A, Jin Y, Tang J-W. 2022. The effect of stand age on fine root production and soil available nutrients in rubber monocultures versus rubber-Flemingia macrophylla agroforestry systems. Forests 13: 830. https://doi.org/10.3390/f13060830   

20. Li Y-P, Feng Y-L, Li W-T, Tomlinson K, Liao Z-Y, Zheng Y-L, Zhang J-L. 2022. Leaf trait association in relation to herbivore defense, drought resistance, and economics in a tropical invasive plant. American Journal of Botany. http://doi.org/10.1002/ajb2.1858 

21. Katuwal HB, Rai J, Tomlinson KW, …, Quan R-C. 2022. Seasonal variation and crop diversification shape the composition of bird communities in agricultural landscapes in Nepal. Agriculture, Ecosystems and Environment. https://doi.org/10.1016/j.agee.2022.107973 

22. Lefebvre T, Charles-Dominique T, Tomlinson KW*. 2022. Trunk spines of trees: a physical defence against bark removal and climbing by mammals? Annals of Botany 129: 541-554. https://doi.org/10.1093/aob/mcac025 

23. Lapuz RS, Monzon AKV, Reyes SR, de Alban JD, Tomlinson KW*. 2022. Greater loss and fragmentation of savannas than forests over the last three decades in Yunnan Province, China. Environmental Research Letters 17: 014003. https://doi.org/10.1088/1748-9326/ac3aa2   

24. Falster D, Gallagher R, …, Tomlinson KW, … 2021. AusTraits, a curated plant trait database for the Australian flora. Scientific Data 8: 254. https://doi.org/10.1038/s41597-021-01006-6 

25. Chu Y, Wee AK, Lapuz RS, Tomlinson KW*. 2021. Phylogeography of two widespread C4 grass species suggest tableland and valley grassy biome in southwestern China pre-date human modification. GECCO, e01835. https://doi.org/10.1016/j.gecco.2021.e01835 

26. Chen Y-J, …, Tomlinson KW, …. 2021. Hydraulic prediction of drought-induced plant dieback and top-kill depends on leaf habit and growth form. Ecology Letters 24: 2350-2363. https://doi.org/10.1111/ele.13856 

27. Maurin M, , Tomlinson KW, . 2021. A nuclear phylogenomic study of the angiosperm order Myrtales, exploring the potential and limitations of the universal Angiosperms353 probe set. American Journal of Botany 108: 1087-1111. https://doi:10.1002/ajb2.1699 

28. Zeeshan M, Sha L, Tomlinson KW, Azeez PA. 2021. Factors shaping students' perception of climate change in the western Himalayas, Jammu & Kashmir, India. Current Research in Environmental Sustainability 3: 100035. https://doi.org/10.1016/j.crsust.2021.100035 

29. Anest A, Charles-Dominique T, Maurin O, Millan M, Edelin C, Tomlinson KW*. 2021. Evolving the structure: climatic and developmental constraints on the evolution of plant architecture. New Phytologist 231: 1278-1295. https://doi.org/10.1111/nph.17296 

30. Harapan TS, Agung AP, Handika H, Novarino W, Tjong DH, Tomlinson KW. 2020. New records and potential geographic distribution of Elongated Caecilian, Ichthyophis elongatus Taylor, 1965 (Amphibia, Gymnophiona, Ichthyophiidae), endemic to West Sumatra, Indonesia. Check List 16: 1695–1701. https://doi.org/10.15560/16.6.1695  

31. Scheiter S, Kumar D, Corlett RT, Gaillard C, Langan L, Lapuz RS, Pfeiffer M, Tomlinson KW. 2020. Climate change promotes transitions to tall evergreen vegetation in tropical Asia. Global Change Biology 26: 5106-5124. https://doi.org/10.1111/gcb.15217 

32. Huang G, Sreekar R, Velho N, Corlett RT, Quan R-C, Tomlinson KW. 2020. Combining camera-trap surveys and interviews to determine the status of mammals in protected rainforests and rubber plantations of Menglun, Xishuangbanna, China. Animal Conservation 23: 689-699. https://doi.org/10.1111/acv.12588 

33. Gao J, Long Z, Zhao W, Wang X-R, Tomlinson KW, Xia S, Chen J. 2020. Combined genotype and phenotype analyses reveal patterns of genomic adaptation to local environments in the subtropical oak Quercus acutissima. Journal of Systematics and Evolution 59: 541-556. https://doi.org/10.1111/jse.12568 

34. Armani M, Goodale UM, Charles-Dominique T, Barton KE, Yao X, Tomlinson KW*. 2020. Structural defence is coupled with the leaf economic spectrum across saplings of spiny species. Oikos 129: 740-752. https://doi.org/10.1111/oik.06960 

35. Corlett RT, Tomlinson KW. 2020. Climate change and edaphic specialists: irresistible force meets immovable object?  Trends in Ecology and Evolution 35: 367-376.  https://doi.org/10.1016/j.tree.2019.12.007 

36. Xu W, Tomlinson KW, Jie L. 2020. Strong intraspecific trait variation in a tropical dominant tree species along an elevational gradient. Plant Diversity 42: 1-6. https://doi.org/10.1016/j.pld.2019.10.004 

37. Yue X, Hughes ACH, Tomlinson KW, Xia S, Li S, Chen J. 2020. Body size and diet–related morphological variation of bats over the past 65 years in China. Journal of Mammalogy 101: 61-79. https://doi.org/10.1093/jmammal/gyz161 

38. Roeder M, Liu JJ, Pasion BOP, Yasuda M, Slik JWF, Tomlinson KW. 20.19. Growth, mortality and wood density of lianas in fragmented forest in SW China. Journal of Vegetation Science 30: 1143-1152. 

39. Armani M, Charles-Dominique T, Barton KE, Tomlinson KW*. 2019. Developmental constraints and resource environment shape early emergence and investment in spines in saplings. Annals of Botany 124: 1133-1142. https://doi.org/10.1093/aob/mcz152 

40. Song H-K, Singh D, Tomlinson KW, Yang X-D, Ogwu M, Slik JWF, Adams JM. 2019. Tropical forest conversion to rubber plantation in southwest China results in lower fungal beta diversity and reduced network complexity. FEMS Microbiology Ecology 95: fiz092. https://doi.org/10.1093/femsec/fiz092 

41. Siddiq Z, Tomlinson KW, Cao KF. 2019. Does fluctuation of meteorological conditions across years influence stand transpiration of Tectona grandis plantation? Ecohydrology 12: e2116. https://doi.org/10.1002/eco.2116 

42. Khaing TT, Pasion BO, Lapuz RS, Tomlinson KW*. 2019. Determinants of plant composition, diversity and structure in a seasonally dry forest in Myanmar. Global Ecology and Conservation 19: e00669. https://doi.org/10.1016/j.gecco.2019.e00669 

43. Roeder M, Yang W, Tomlinson KW. 2019. Influence of smoke, heat and fire on germination of woody species occurring in the dry valleys of southwest China. Journal of Plant Ecology 12: 931-940. https://doi.org/10.1093/jpe/rtz030  (Editor’s choice) 

44. Singh D, Slik FJW, Jeon YS, Tomlinson KW, Yang X-D, Wang J, Porazinska DL, Adams JM. 2019. Tropical forest conversion to rubber plantation affects soil micro- & mesofaunal community & diversity. Scientific Reports 9: 5893. https://www.nature.com/articles/s41598-019-42333-4 

45. Tomlinson KW*, Sterck FJ, Barbosa ERM, de Bie S, Prins HHT, van Langevelde F. 2019. Seedling growth of savanna tree species from three continents under grass competition and nutrient limitation in a greenhouse experiment. Journal of Ecology 107: 1051-1066. https://doi.org/10.1111/1365-2745.13085 

46. Armani M, van Langevelde F, Tomlinson KW, Adu-Bredu S, Djagbletey G, Veenendaal EM. 2018. Compositional patterns of overstorey and understorey woody communities in a forest-savanna boundary in Ghana. Plant Ecology & Diversity 11: 451-463. https://doi.org/10.1080/17550874.2018.1539133 

47. Pasion BO, Roeder M, Liu J-J, Yasuda M, Corlett RT, Slik JFW, Tomlinson KW*. 2018. Trees represent community composition of other plant life-forms, but not their diversity, abundance or responses to fragmentation. Scientific Reports 8: 11374. https://www.nature.com/articles/s41598-018-29635-9 

48. Beng KC, Corlett RC, Tomlinson KW. 2018. Seasonal changes in the diversity and composition of the litter fauna in natural forests and rubber plantations. Scientific Reports 8: 10232. https://doi.org/10.1038/s41598-018-28603-7 

49. Charles-Dominique T, Midgley GF, Tomlinson KW, Bond WJ. 2018. Steal the light: Shade versus fire adapted vegetation in forest-savanna mosaics. New Phytologist 218: 1419-1429. https://doi.org/10.1111/nph.15117 

50. Wang B, Phillips JS, Tomlinson KW. 2018. Trade-off between physical and chemical defence in plant seeds is mediated by seed mass. Oikos 127: 440-447. https://doi.org/10.1111/oik.04867 

51. Griffith DM, …, Tomlinson KW, …. 2017.  Comment on “The extent of forest in dryland biomes”. Science 358: eaao1309. http://doi.org/10.1126/science.aao1309 

52. Dayananda S, …, Tomlinson K, Goodale E. 2017. Topography and soil type are critical to understanding how bird and herpetofaunal communities persist in forest fragments of tropical China. Biological Conservation 215: 107-115. https://www.sciencedirect.com/science/article/pii/S0006320717300526.  

53. Padmanaba M, Tomlinson KW, Hughes ACH, Corlett RT. 2017. Alien plant invasions of protected areas in Java, Indonesia. Scientific Reports 7: 9334. https://doi.org/10.1038/s41598-017-09768-z 

54. Chen Q, Tomlinson KW, Cao L, Wang B. 2017. Effects of fragmentation on the seed predation and dispersal by rodents differ among species with different seed size. Integrative Zoology 12: 468-476. http://onlinelibrary.wiley.com/doi/10.1111/1749-4877.12273/full  

55. Liu J-J, Yu M-J, Tomlinson K, Slik JWF 2017. Patterns and drivers of plant biodiversity in Chinese university campuses. Landscape and Urban Planning 164: 64-70. https://doi.org/10.1016/j.landurbplan.2017.04.008 

56. Chaturvedi RK, Raghubanshi AS, Tomlinson KW & Singh JS. 2017. Impacts of human disturbance in tropical dry forests increase with soil moisture stress. Journal of Vegetation Science 28: 997-1007. http://onlinelibrary.wiley.com/doi/10.1111/jvs.12547/full.  

57. Leles B, Xiao X, Pasion B, Nakamura A, Tomlinson KW. 2017. Does plant diversity increase top-down control of herbivorous insects in tropical forest? Oikos 126: 1142-1149. http://onlinelibrary.wiley.com/doi/10.1111/oik.03562/full  

58. Ratnam J, Tomlinson KW*, Rasquinha DN & Sankaran M. 2016. Savannahs in Asia: evidence for antiquity, biogeography, and an uncertain future. Philosophical Transactions of the Royal Society B 371: 20150305 http://dx.doi.org/10.1098/rstb.2015.0305. (Joint First Author) 

59. Chen Y-J, Schnitzer S; Zhang Y-J, Fan Z-X, Goldstein G, Tomlinson KW, Cao K-F, Zhang J-L 2017. Physiological regulation and efficient xylem water transport regulate diurnal water and carbon balances of tropical lianas. Functional Ecology 31: 306-317. https://doi.org/10.1111/1365-2435.12724.   

60. Dayananda SK, Goodale E, Lee M-B, Liu J-J, Mammides C, Pasion BO, Quan R-C, Slik JWF, Sreekar R, Tomlinson KW, Yasuda M. 2016. Effects of forest fragmentation on nocturnal Asian birds: a case study from Xishuangbanna, China. Zoological Research 37: 151-158. https://doi.org/10.13918/j.issn.2095-8137.2016.3.151 

61. Beng KC, Tomlinson KW, Shen XH, Surget-Groba Y, Hughes AC, Corlett RT, Slik JWF. 2016. The utility of DNA metabarcoding for studying the response of arthropod diversity and composition to land-use change in the tropics. Scientific Reports 6: 24965.  http://www.nature.com/articles/srep24965  

62. Monoy CC, Tomlinson KW*, Iida Y, Swenson N, Slik FJW. 2016. Temporal changes in tree species and trait composition in a cyclone-prone Pacific dipterocarp forest. Ecosystems 19: 1013–1022.  https://doi.org/10.1007/s10021-016-9983-0 

63. Sreekar R, Huang G, Yasuda M, Quan R-C, Goodale E, Corlett R, Tomlinson KW. 2016. Effects of forests, roads and mistletoe on bird diversity in monoculture rubber plantations. Scientific Reports 6: 21822.    http://www.nature.com/articles/srep21822 

64. Tomlinson KW*, van Langevelde F, Ward DM, Prins HHT de Bie S, Vosman B, Sampaio EVSB & Sterck F. 2016. Defence against vertebrate herbivores diverges into architectural and low nutrient strategies amongst savanna tree species. Oikos 125: 126-136. https://doi.org/10.1111/oik.02325 

65. Bhatt, P, Rawat GS, Sankar K, Tomlinson K, van Langevelde F. 2015. Pattern of invasion by Adhatoda vasica in savannahs of Sariska Tiger Reserve, Rajasthan, western India. Pp 170 – 179. In: Rawat GS, Adhikari BS (Eds.) Ecology and Management of Grassland Habitats in India, ENVIS Bulletin. Wildlife and Protected Areas. WII Dehradun. [BOOK CHAPTER] 

66. Chen Y-J, Bongers F, Tomlinson K, Fan Z-X, Lin H, Zhang S-B, Zheng Y-L, Li Y-P, Cao K-F, Zhang J-L. 2015. Time lags between crown and basal sap flows in tropical lianas and co-occurring trees. Tree Physiology 36: 736-747. doi:10.1093/treephys/tpv103 

67. Roeder M, Slik FJW, Harrison RD, Paudel E, Tomlinson KW. 2015. Proximity to the host is an important characteristic for the selection of the first support in lianas. Journal of Vegetation Science 26: 1054-1060. 

68. Tomlinson KW*, Poorter L, Bongers F, Borghetti F, Jacobs L, van Langevelde F. 2014. Relative growth rate variation of evergreen and deciduous savanna tree species is driven by different traits. Annals of Botany 114: 315-324.  

69. Tomlinson KW*. 2014. The mixed success of Mimosoideae clades invading into Australia. In: Prins HHT, Gordon I (eds) Invasion biology and ecological theory: insights from a continent in transformation. Cambridge University Press, Cambridge, UK. Pp 39-57. [BOOK CHAPTER] 

70. Barbosa ERM, Tomlinson KW, Carvalheiro LG, Kirkman K, de Bie S, Prins HHT, van Langevelde F. 2014. Short-term effect of nutrient availability and rainfall distribution on biomass production and leaf nutrient content of savanna tree species. PLoS ONE 9: e92619. doi:10.1371/journal.pone.0092619 

71. Barbosa ERM, van Langevelde F, Tomlinson KW, Kirkman K, de Bie S, Prins HHT. 2014. Tree species from different functional groups respond differently to environmental changes during establishment.  Oecologia 174: 1345-1357. doi:10.1007/s00442-013-2853-y 

72. Tomlinson KW*, van Langevelde F, Ward D, Bongers F, da Silva DA, Prins HHT, de Bie S & Sterck FJ. 2013. Deciduous and evergreen trees differ in juvenile biomass allometries because of differences in allocation to root storage. Annals of Botany 112: 575–587. doi:10.1093/aob/mct132 

73. Tomlinson KW*, Poorter L, Sterck FJ, Borghetti F, Ward D, de Bie S & van Langevelde F 2013. Leaf adaptations of evergreen and deciduous trees of semi-arid and humid savannas on three continents.  Journal of Ecology 101: 430-440. doi: 10.1111/1365-2745.12056.x 

74. Tomlinson KW*, Sterck FJ, Bongers F, da Silva DA, Barbosa ERM, Ward D, et al. 2012. Biomass partitioning and root morphology of savanna trees across a water gradient. Journal of Ecology 100: 1113-1121. https://doi.org/10.1111/j.1365-2745.2012.01975.x 

75. Moore S, Shrestha S, Tomlinson KW & Vuong H. 2011. Predicting the effect of climate change on African trypanosomiasis: integrating epidemiology with parasite and vector biology. Journal of the Royal Society Interface 7: 22072451 doi: 10.1098/rsif.2011.0654.   

76. Van Langevelde F, Tomlinson KW, Barbosa ERM, de Bie S, Prins HHT, Higgins SI 2011. Understanding tree-grass coexistence and impacts of disturbance and resource variability in savannas. In: Hill MJ, Hanan NL (eds). Ecosystem function in savannas: measurement and modelling at landscape to global scales. CRC Press, Boca Raton, USA. pp 257-271.  [BOOK CHAPTER] 

77. Jewitt GPW, …, Tomlinson K. 2009. Methods and guidelines for the licensing of SFRAs with particular reference to low flows. Water Research Commission.  Report No. 1428/1/09. Pretoria, South Africa. [REPORT] 

78. Tomlinson KW*, Hearne JW, Dominy JG & O’Connor TG 2007. A functional-structural model for growth of clonal bunchgrasses. Ecological Modelling 202: 243-264. 

79. Tomlinson KW* & O’Connor TG 2005. The effect of defoliation environment on primary growth allocation and secondary tiller recruitment of two bunchgrasses. African Journal of Range and Forage Science 22: 29-36. 

80. Tomlinson KW* & O’Connor TG 2004. Control of tiller recruitment in bunchgrasses: uniting physiology and ecology. Functional Ecology 18: 489-496. 

81. Tomlinson KW, Hearne JW & Alexander RR 2002. An approach to evaluate the effect of property size on land-use options in semi-arid rangelands. Ecological Modelling 149: 85-95. 





1. Savanna Science Networking Meeting. 1 - 5 March 2020 Skukuza, South Africa. Oral presentation. 

2. Gordon Research Conference on plant-herbivore interactions. 24 February – 1 March 2019, Ventura, California, USA. Poster. 

3. Meeting of the Ecological Society of Yunnan, 4-5 December 2017, XTBG, China. Invited Plenary Lecture. 

4. 5th Workshop of Science and Technology Cooperation. Joint CAS-TRF meeting, 6 – 8 November 2017, Chiang Mai, Thailand. Poster. 

Community service:

Commitment to research the situation:



2023 Yunnan Province Foundational Research Key Grant (云南省基础研究专项-重点项目), 202301AS070238, Amount: 500,000 RMB. 

Project title: Impacts of forest fragment configuration on genetic diversity and population viability of tree species with different disperser guilds in defaunated landscapes 


2020 Yunnan Province Thousand Talents Plan, E0YN021. Amount: 1,000,000 RMB. 

Project title: The role of domestic herbivory in shaping landscapes in Yunnan. 


2018 National Natural Science Foundation of China (NSFC), 31870425, Amount: 600,000 RMB. 

Project title: Biogeography and biodiversity of Southeast Asian pine savannas and forests under past and future climates. 


2015 Joint NSFC-Yunnan Government Key Grant, U1502264. Amount: 2,300,736 RMB. 

Project title:  Geography, diversity and conservation of Savanna Ecosystems in Yunnan, China. 


2014 National Natural Science Foundation of China (NSFC), 31470449, Amount: 840,000 RMB.  

Project title: Ecology and distribution of spinescent plants in Yunnan.