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Faculty and Staff
QUAN Ruichang
Deputy Director
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Mailing Address:
Center for Integrative Conservation, XTBG, Xishuangbanna, Yunnan 666303,P. R. China   Tel: 86-691-8713156



  Personal information 

  Name: Quan Ruichang      

  Birth date: January 28, 1973      Gender: Male 

  Nationality: P. R. China         


  9/1998-7/2003, Ph.D. Course, Ph.D. Kunming Institute of Zoology (KIZ), Chinese Academy of Science 

  9/1994-7/1998, Bachelor Course, Bachelor. University of Southwest Forestry.  

  Professional experiences 

  1/2013-Present: Professor (XTBG) 

  8/2007-12/2012: Associated Professor (XTBG) 

  1/2005-7/2007: Assistant Professor (KIZ)  

  Major research directions: animal/behavioral ecology, biodiversity conservation (particularly birds and mammals), animal-mediated seed dispersal (birds and mammals). 





1.Than KZ, Zaw Z, Quan RC, Hughes AC: Biodiversity conservation in Myanmar's coastal wetlands: Focusing on saltwater crocodile habitats and connectivity. Biological Conservation 2024, 289. 

2.Katuwal HB, Sharma HP, Rokka P, Das NK, Quan RC: Knowledge, attitudes, and conservation challenges for the Lesser Adjutant in Nepal. Global Ecology and Conservation 2024, 49. 

3.Hua FY, Wang WY, Nakagawa S, Liu SQ, Miao XR, Yu L, Du ZR, Abrahamczyk S, Arias-Sosa LA, Buda K et al: Ecological filtering shapes the impacts of agricultural deforestation on biodiversity. Nature Ecology & Evolution 2024, 8(2):188-189. 

4.Thu AM, Thang TH, Campos-Arceiz A, Quan RC: Disentangling vertebrate spatio-temporal responses to anthropogenic disturbances: evidence from a protected area in central Myanmar. Oryx 2023, 57(4):512-521. 

5.Thu AM, Phyo AM, Quan RC: First camera-trap evidence of banded civet<i> Hemigalus derbyanus </i>(Mammalia: Carnivora: Viverridae) in Myanmar. Mammalia 2023, 87(2):158-161. 

6.Lwin YH, Wang L, Li GG, Quan RC: Where do we manage? Uncovering the hidden impact of human presence on mammal conservation in northern Myanmar. Biological Conservation 2023, 284. 

7.Katuwal HB, Sharma HP, Thakur RK, Rokka P, Mandal DN, Baral HS, Quan RC: Illegal trapping and local trade of farmland birds in Madhesh Province, Nepal. Global Ecology and Conservation 2023, 42. 

8.Katuwal HB, Sharma HP, Rokka P, Bhusal KP, Bhattarai BP, Koirala S, Luitel SC, Yadav S, Sah G, Baral HS et al: The effects of climate and land use change on the potential distribution and nesting habitat of the Lesser Adjutant in Nepal. Avian Research 2023, 14. 

9.He RC, Wang L, Goodale E, Li K, Quan RC: The extremely small body size of Williamson's mouse deer (Tragulus williamsoni) allows coexistence with sympatric larger ungulates through temporal avoidance. Frontiers in Ecology and Evolution 2023, 11. 

10.Yu WH, Lin CY, Huang ZLY, Liu S, Wang QY, Quan RC, Li S, Wu Y: Discovery of Kerivoula kachinensis and a validity of K. titania (Chiroptera: Vespertilionidae) in China. Mammalia 2022, 86(3):303-308. 

11.Liu S, Rao DQ, Zhang DR, Lwin YH, Mo MZ, Zuo CS, Yin FW, Quan RC, Li S: Phylogenetic position of Odorrana macrotympana (Yang, 2008) (Anura, Ranidae) and extension of its geographical distribution. Herpetozoa 2022, 35:9-19. 

12.Li LL, He RC, Pansini R, Quan RC: Prolonged proximity to humans ensures better performance of semi-captive Asian elephants at discriminating between human individuals by voice. Frontiers in Ecology and Evolution 2022, 10. 

13.Katuwal HB, Sundar KSG, Zhang MX, Rimal B, Baral HS, Sharma HP, Ghimire P, Hughes AC, Quan RC: Factors affecting the breeding ecology of the globally threatened Lesser Adjutant<i> (Leptoptilosjavanicus)</i> in agricultural landscapes of Nepal. Avian Conservation and Ecology 2022, 17(2). 

14.Katuwal HB, Sundar KSG, Zhang MX, Rimal B, Baral HS, Sharma HP, Ghimire P, Hughes AC, Quan RC: Factors affecting the breeding ecology of the globally threatened Lesser Adjutant (Leptoptilosjavanicus) in agricultural landscapes of Nepal. Avian Conservation and Ecology 2022, 17(2). 

15.Katuwal HB, Rai J, Tomlinson K, Rimal B, Sharma HP, Baral HS, Hughes AC, Quan RC: Seasonal variation and crop diversity shape the composition of bird communities in agricultural landscapes in Nepal. Agriculture Ecosystems & Environment 2022, 333. 

16.Dong B, Liu JX, Quan RC, Chen ZQ: Mirror image stimulation could reverse social-isolation-induced aggressiveness in the high-level subsocial lactating spider. Anim Cogn 2022, 25(5):1345-1355. 

17.Chen C, Brodie JF, Kays R, Davies TJ, Liu RZ, Fisher JT, Ahumada J, McShea W, Sheil D, Agwanda B et al: Global camera trap synthesis highlights the importance of protected areas in maintaining mammal diversity. Conservation Letters 2022, 15(2). 

18.Zhou LP, Peabotuwage I, Luo K, Quan RC, Goodale E: Using playback to test leadership in mixed-species flocks and compare flocking with mobbing. Animal Behaviour 2021, 180:151-166. 

19.Zhang YC, Lwin YH, Li R, Maung KW, Li GG, Quan RC: Molecular phylogeny of the genus Muntiacus with special emphasis on the phylogenetic position of Muntiacus gongshanensis. Zoological Research 2021, 42(2):212-216. 

20.Yu WH, Csorba G, Huang ZLY, Li YN, Liu S, Quan RC, Wang QY, Shi HY, Wu Y, Li S: First record of disk-footed bat Eudiscopus denticulus (Chiroptera, Vespertilionidae) from China and resolution of phylogenetic position of the genus. Zoological Research 2021, 42(1):94-99. 

21.Yan MX, Gu BJ, Zhang MX, Wang W, Quan RC, Li JQ, Wang L: The Range Contraction and Future Conservation of Green Peafowl (Pavo muticus) in China. Sustainability 2021, 13(21). 

22.Wang L, Yang B, Bai Y, Lu XQ, Corlett RT, Tan YH, Chen XY, Zhu JG, Liu Y, Quan RC: Conservation planning on China's borders with Myanmar, Laos, and Vietnam. Conservation Biology 2021, 35(6):1797-1808. 

23.Thein MM, Wu LM, Corlett RT, Quan RC, Wang B: Changes in seed predation along a 2300-m elevational gradient on a tropical mountain in Myanmar: a standardized test with 32 non-native plant species. Ecography 2021, 44(4):602-611. 

24.Meaux E, Peabotuwage I, Mammides C, Malykhina K, Quan RC, Goodale E: Behavioural variables influence contact call rate more than characteristics of the vegetation in a group-living passerine species. Behavioural Processes 2021, 185. 

25.Lwin YH, Wang L, Li GG, Maung KW, Swa K, Quan RC: Diversity, distribution and conservation of large mammals in northern Myanmar. Global Ecology and Conservation 2021, 29. 

26.Liu CY, Ye P, Cai Y, Quan RC, Yang CC: Persistent fine-tuning of egg rejection based on parasitic timing in a cuckoo host even after relaxation of parasitism pressure. Behavioural Processes 2021, 193. 

27.Liu CY, Gelin U, He RC, Li H, Quan RC: Flexible breeding performance under unstable climatic conditions in a tropical passerine in Southwest China. Zoological Research 2021, 42(2):221-226. 

28.Li LL, Plotnik JM, Xia SW, Meaux E, Quan RC: Cooperating elephants mitigate competition until the stakes get too high. Plos Biology 2021, 19(9). 

29.Katuwal HB, Zhang MX, Baral HS, Sharma HP, Quan RC: Assessment of farmers' knowledge and perceptions towards farmland birds show the need of conservation interventions. Global Ecology and Conservation 2021, 27. 

30.Chen S, Sun GZ, Wang Y, Huang C, Chen Y, Liu P, Deng Y, Cao DF, Zhang MX, Ong S et al: A multistakeholder exercise to identify research and conservation priorities for Asian elephants in China. Global Ecology and Conservation 2021, 27. 

31.Zhou SS, Quan RC, Li R, Liu Q, Yin JT: Colocasia kachinensis, a new species of Araceae from Myanmar. Phytokeys 2020(138):41-47. 

32.Zhou SS, Li R, Quan RC, Shine L, Duan LD: Ophiorrhiza monsvictoriae (Rubiaceae, Rubioideae), a new species from Myanmar. Phytokeys 2020(138):219-223. 

33.Sun N, Cao GH, Li GG, Liu ZH, Quan RC: Macaca leonina has a wider niche breadth than sympatric M. mulatta in a fragmented tropical forest in southwest China. American Journal of Primatology 2020, 82(2). 

34.Luo K, Xu ZD, Wang X, Quan RC, Lu ZY, Bi WQ, Zhao H, Qiu GL: Terrestrial methylmercury bioaccumulation in a pine forest food chain revealed by live nest videography observations and nitrogen isotopes. Environmental Pollution 2020, 263. 

35.Liu S, Lwin YH, Quan RC, Li S: First record of Rhacophorus verrucopus Huang, 1983 from Myanmar. Herpetozoa 2020, 33:207-211. 

36.Li GG, Sun N, Swa K, Zhang MX, Lwin YH, Quan RC: Phylogenetic reassessment of gorals with new evidence from northern Myanmar reveals five distinct species. Mammal Review 2020, 50(4):325-330. 

37.Li GG, Lwin YH, Yang B, Qin T, Phothisath P, Maung KW, Quan RC, Li S: Taxonomic revision and phylogenetic position of the flying squirrel genus Biswamoyopterus (Mammalia, Rodentia, Sciuridae, Pteromyini) on the northern Indo-China peninsula. Zookeys 2020(939):65-85. 

38.Gan YL, Qin T, Lwin YH, Li GG, Quan RC, Liu S, Yu GH: A new species of Amolops (Anura: Ranidae) from northern Myanmar. Zoological Research 2020, 41(6):733-+. 

39.Cao YL, Li R, Zhou SS, Song L, Quan RC, Hu HB: Ethnobotanical study on wild edible plants used by three trans-boundary ethnic groups in Jiangcheng County, Pu'er, Southwest China. Journal of Ethnobiology and Ethnomedicine 2020, 16(1). 

40.Aung MH, Li DZ, Tan YH, Xia NH, Quan RC, Jin XH: Documentation of plant diversity of Southeast Asia: the new role of Belt and Road Initiative. Phytokeys 2020(138):1-2. 

41.Zhou LP, Peabotuwage I, Gu H, Jiang DM, Hu GH, Jiang AW, Mammides C, Zhang MX, Quan RC, Goodale E: The response of mixed-species bird flocks to anthropogenic disturbance and elevational variation in southwest China. Condor 2019, 121(3). 

42.Zhang YC, Chen XY, Li GG, Quan RC: Complete mitochondrial genome of Gongshan muntjac ( Muntiacus gongshanensis), a Critically Endangered deer species. Mitochondrial DNA Part B-Resources 2019, 4(2):2867-2868. 

43.Thu AM, Li GG, Zhang MX, Thang TH, Soe AM, Naing W, Quan RC: Group size and social organization of the endangered Eld's deer (Rucervus eldii thamin): Results from a long-term study in Myanmar. Global Ecology and Conservation 2019, 18. 

44.Luo K, Feng L, Lu ZY, Li DW, Quan RC: Novel instance of brood parasitic cuckoo nestlings using bright yellow patches to mimic gapes of host nestlings. Wilson Journal of Ornithology 2019, 131(3):686-+. 

45.Li L, Goodale E, Quan RC: Nest predation on an abundant generalist bird in tropical China. Wilson Journal of Ornithology 2019, 131(3):514-523. 

46.Dong B, Quan RC, Chen ZQ: Prolonged milk provisioning and extended maternal care in the milking spider Toxeus magnus: biological implications and questions unresolved. Zoological Research 2019, 40(4):241-243. 

47.Chen C, Quan RC, Cao GH, Yang HP, Burton AC, Meitner M, Brodie JF: Effects of law enforcement and community outreach on mammal diversity in a biodiversity hotspot. Conservation Biology 2019, 33(3):612-622. 

48.Chang CH, Williams SJ, Zhang MX, Levin SA, Wilcove DS, Quan RC: Perceived entertainment and recreational value motivate illegal hunting in Southwest China. Biological Conservation 2019, 234:100-106. 

49.Zhou SS, Tan YH, Li R, Quan RC, Maung KW, Liu Q, Sima YK: Magnolia kachinensis (Magnoliaceae), a new species from northern Myanmar. Phytotaxa 2018, 375(1):92-98. 

50.Zhou SS, Tan YH, Jin XH, Maung KW, Zyaw M, Li R, Quan RC, Liu Q: Coelogyne victoria-reginae (Orchidaceae, Epidendroideae, Arethuseae), a new species from Chin State, Myanmar. Phytokeys 2018(98):125-133. 

51.Liu Q, Zhou SS, Jin XH, Pan B, Maung KW, Zyaw M, Li R, Quan RC, Tan YH: Dendrobium naungmungense (Orchidaceae, Dendrobieae), a new species from Kachin State, Myanmar. Phytokeys 2018(94):31-38. 

52.Jin XH, Tan YH, Quan RC: Taxonomic discoveries bridging the gap between our knowledge and biodiversity. Phytokeys 2018(94):1-2. 

53.Chen ZQ, Corlett RT, Jiao XG, Liu SJ, Charles-Dominique T, Zhang SC, Li H, Lai R, Long CB, Quan RC: Prolonged milk provisioning in a jumping spider. Science 2018, 362(6418):1052-+. 

54.Zhang MX, Kyaw M, Li GG, Zhao JB, Zeng XL, Swa K, Quan RC: Bird diversity in northern Myanmar and conservation implications. Zoological Research 2017, 38(5):264-280. 

55.Zhang MX, Gouveia A, Qin T, Quan RC, Nijman V: Illegal pangolin trade in northernmost Myanmar and its links to India and China. Global Ecology and Conservation 2017, 10:23-31. 

56.Zhang MX, Chang C, Quan RC: Natural forest at landscape scale is most important for bird conservation in rubber plantation. Biological Conservation 2017, 210:243-252. 

57.Yang B, Zhou SS, Liu Q, Maung KW, Li R, Quan RC, Tan YH: Coelogyne magnifica (Orchidaceae), a new species from northern Myanmar. Phytokeys 2017(88):109-117. 

58.Liu Q, Zhou SS, Li R, Zhang MX, Zyaw M, Lone S, Quan RC: Bulbophyllum putaoensis (Orchidaceae: Epidendroideae; Malaxideae), a new species from Kachin State, Myanmar. Phytotaxa 2017, 305(1):57-60. 

59.Li SQ, Quan RC: Taxonomy is the cornerstone of biodiversity conservation - SEABRI reports on biological surveys in Southeast Asia. Zoological Research 2017, 38(5):213-214. 

60.Li GG, Zhang MX, Swa K, Maung KW, Quan RC: Complete mitochondrial genome of the leaf muntjac (Muntiacus putaoensis) and phylogenetics of the genus Muntiacus. Zoological Research 2017, 38(5):310-316. 

61.Guo W, Cao GH, Quan RC: Population dynamics and space use of wild boar in a tropical forest, Southwest China. Global Ecology and Conservation 2017, 11:115-124. 

62.Dayananda SK, Mammides C, Lee MB, Liu JJ, Pasion BO, Sreekar R, Yasuda M, Quan RC, Slik JWF, Tomlinson KW et al: Topography and soil type are critical to understanding how bird and herpetofaunal communities persist in forest fragments of tropical China. Biological Conservation 2017, 215:107-115. 

63.Chang CH, Barnes ML, Frye M, Zhang MX, Quan RC, Reisman LMG, Levin SA, Wilcove DS: The pleasure of pursuit: recreational hunters in rural Southwest China exhibit low exit rates in response to declining catch. Ecology and Society 2017, 22(1). 

64.Abeysinghe KS, Yang XD, Goodale E, Anderson CWN, Bishop K, Cao AX, Feng XB, Liu SJ, Mammides C, Meng B et al: TOTAL MERCURY AND METHYLMERCURY CONCENTRATIONS OVER A GRADIENT OF CONTAMINATION IN EARTHWORMS LIVING IN RICE PADDY SOIL. Environmental Toxicology and Chemistry 2017, 36(5):1202-1210. 

65.Abeysinghe KS, Qiu GL, Goodale E, Anderson CWN, Bishop K, Evers DC, Goodale MW, Hintelmann H, Liu SJ, Mammides C et al: Mercury flow through an Asian rice-based food web. Environmental Pollution 2017, 229:219-228. 

66.Sreekar R, Huang G, Yasuda M, Quan RC, Goodale E, Corlett RT, Tomlinson KW: Effects of forests, roads and mistletoe on bird diversity in monoculture rubber plantations. Scientific Reports 2016, 6. 

67.Dayananda SK, Goodale E, Lee MB, Liu JJ, Mammides C, Pasion BO, Quan RC, Slik JWF, Sreekar R, Tomlinson KW et al: Effects of forest fragmentation on nocturnal Asian birds: A case study from Xishuangbanna, China. Zoological Research 2016, 37(3):151-158. 

68.Sreekar R, Huang GHL, Zhao JB, Pasion BO, Yasuda M, Zhang K, Peabotuwage I, Wang X, Quan RC, Slik JWF et al: The use of species-area relationships to partition the effects of hunting and deforestation on bird extirpations in a fragmented landscape. Diversity and Distributions 2015, 21(4):441-450. 

69.Quan RC, Li H: Nest Defense by a White-rumped Shama (Copsychus malabaricus) Against Snakes. Wilson Journal of Ornithology 2015, 127(3):538-542. 

70.Duan Q, Goodale E, Quan RC: Bird fruit preferences match the frequency of fruit colours in tropical Asia. Scientific Reports 2014, 4. 

71.Wan Y, Quan RC, Ren GP, Wang L, Long YC, Liu XH, Zhu JG: Niche Divergence Among Sex and Age Classes in Black-and-White Snub-nosed Monkeys (Rhinopithecus bieti). International Journal of Primatology 2013, 34(5):946-956. 

72.Quan RC, Cui LW: Winter pairs of ruddy shelducks at Lashihai Lake, southwest China. Integrative Zoology 2013, 8:46-49. 


74.Chang X, Quan RC, Wang L: Bird conservation in extremely small tropical rainforest patches in southwest China. Biological Conservation 2013, 158:188-195. 

75.Quan RC, Ren GP, Behm JE, Wang L, Huang Y, Long YC, Zhu JG: Why Does Rhinopithecus bieti Prefer the Highest Elevation Range in Winter? A Test of the Sunshine Hypothesis. Plos One 2011, 6(9). 

76.Quan RC, Huang Y, Warren MW, Zhao QK, Ren GP, Huo S, Long YC, Zhu JG: How Human Household Size Affects the Habitat of Black-and-White Snub-Nosed Monkeys (Rhinopithecus bieti) in Hongla Snow Mountain Nature Reserve in Tibet, China. International Journal of Primatology 2011, 32(5):1190-1202. 

77.Cui LW, Huo S, Zhong T, Xiang ZF, Xiao W, Quan RC: Social organization of black-and-white snub-nosed monkeys (Rhinopithecus bieti) at Deqin, China. American Journal of Primatology 2008, 70(2):169-174. 

78.Xiang ZF, Huo S, Xiao W, Quan RC, Grueter CC: Diet and feeding behavior of Rhinopithecus bieti at Xiaochangdu, Tibet: Adaptations to a marginal environment. American Journal of Primatology 2007, 69(10):1141-1158. 

79.Xiang ZF, Huo S, Wang L, Cui LW, Xiao W, Quan RC, Tai Z: Distribution, status and conservation of the black-and-white snub-nosed monkey Rhinopithecus bieti in Tibet. Oryx 2007, 41(4):525-531. 

80.Cui LW, Quan RC, Xiao W: Sleeping sites of black-and-white snub-nosed monkeys (Rhinopithecus bieti) at Baima Snow Mountain, China. Journal of Zoology 2006, 270(1):192-198. 

81.Quan RC, Cui LW, Wen XJ: Sexual differences in vigilance of paired Ruddy Shelduck in winter. Waterbirds 2003, 26(3):335-337. 

82.Quan RC, Wen XJ, Yang XJ: Numbers of migratory waterbirds at Lake Lashihai, China. Waterbirds 2002, 25(2):239-244. 

83.Quan RC, Wen XJ, Yang XJ: Effects of human activities on migratory waterbirds at Lashihai Lake, China. Biological Conservation 2002, 108(3):273-279. 

84.Quan RC, Wen XJ, Yang XJ, Peng GH, Huang TF: Habitat use by wintering Ruddy Shelduck at Lashihai Lake, Lijiang, China. Waterbirds 2001, 24(3):402-406. 



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